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Life cycle of a predaceous ground beetle (Carabidae). All stages commonly occur only on the ground. Egg laying and pupation occur in litter or the soil surface. Adults of some species climb shrubs or trees to feed on caterpillars.

Flint ML, Dreistadt SH. 1998. Natural Enemies Handbook: The Illustrated Guide to Biological Pest Control. UC ANR Publ. 3386. Oakland, CA. Adapted from adult and pupa by Packard AS. 1872. Guide to the Study of Insects. 3rd. ed. F.W. Putnam. Salem, MA. Larvae by Peterson A. 1960. Larvae of Insects: An Introduction to Nearctic Species. Part II. Edwards Brothers. Ann Arbor, MI. Reprinted with permission of Helen Peterson.

Statewide IPM Program, Agriculture and Natural Resources, University of California
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